David Harmon
Regular Faculty
Professional Biography
Professional Biography
Beef Nutrition - Research/Instruction
Professional Experience
Department of Animal Sciences, University of Kentucky
2002 - present, Director of Graduate Studies
July 1995 - present, Professor
January 1992 - July 1995, Associate Professor
Appointment .75 research - .25 Teaching
February 1989 - December 1991, Associate Professor
Department of Animal Science and Industry, Kansas State University
August 1983 -February 1989, Assistant Professor
Appointment .6 Research - .4 Teaching
Teaching
Experience with graduate and undergraduate
- Companion Animal Nutrition, undergraduate nutrition
- Fundamentals of Nutrition, Introductory undergraduate nutrition.
- Animal Nutrition, Graduate, nutritional physiology and metabolism.
- Analytical Techniques in Animal Science, Graduate laboratory methods.
- Radioisotopes and Radioimmunoassay, Graduate laboratory methods.
- Techniques in Animal Nutrition Research, Graduate lecture and laboratory.
- Advanced Ruminant Nutrition, Graduate lecture.
Research
Research program has dealt with identifying relationships between diet digestion, gut metabolism and nutrient absorption in ruminants and companion animals.
- Principal Investigator on over $4.3 million in research funding
- Co-Investigator on an additional $792,000
Students
Supervised the thesis for 19 M.S. and 15 Ph.D. students who have completed their degrees as well as 7 post-doctoral students.
Additional Activities
Specialty Chief Editor Animal Nutrition, Frontiers in Animal Science (2020-2023)
Member of University Animal Care Use and Committee (KSU and UK)
IACUC College of Agriculture Subcommittee Chair (1997-2004)
Affiliations and Honors
University of Kentucky Research Professor, 2024-2025
William B. Sturgill Award- UK outstanding contributions to graduate education, 2024
Recognized as one of 143 University of Kentucky scientists and scholars that rank among top 2% of world’s most-cited researchers – 2023
American Society of Animal Science Industry Service Award- 2021
American Society of Animal Science Research Fellow Award- 2012
University of Kentucky College of Agriculture George E. Mitchell, Jr. Award for Outstanding Service to Graduate Students- 2010
University of Kentucky College of Agriculture Thomas Poe Cooper Award for Research- 2006
American Society of Animal Science Jim Corbin Award in Companion Animal Biology- 2005
American Society of Animal Science Director at Large 1999-2002
American Society of Animal Science AFIA Ruminant Nutrition Award- 1998
Journal of Animal Science-Section Editor Ruminant Digestion and Metabolism - 1992-1995
Midwest American Society of Animal Science Young Researcher Award - 1992
Finalist- Elanco Young Animal Scientist Research Award, 1991
Chair - 31st annual Ruminant Nutrition Symposium FASEB, 1990, Washington, D.C.
American Society for Nutritional Sciences
American Society of Animal Science
Gamma Sigma Delta
Sigma Xi
Widaman Trust Outstanding Graduate Student Award, 1982.
Scholars@UK
Association of serotonin and ergot alkaloids on tissue partitioning and contractile response of bovine blood vessels
Valente, E. E. L., Harmon, D. L., May, J., Ji, H., Trotta, R. J. & Klotz, J. L., Jan 2026, In: Current Research in Toxicology. 10, 100272.Research output: Contribution to journal › Article › peer-review
Determining the influence of rearing method on lamb weaning behaviour
Simeonov, M. & Harmon, D. L., Oct 10 2025, In: Acta Agriculturae Slovenica. 121, 3Research output: Contribution to journal › Article › peer-review
The effect of weaning methods on stress and the welfare in early weaned lambs
Simeonov, M., Harmon, D. L., Stoycheva, I. & Mihailova, M., Nov 2025, In: Tropical Animal Health and Production. 57, 8, 441.Research output: Contribution to journal › Article › peer-review
Effects of Saccharomyces cerevisiae fermentation-derived postbiotic on methane production and plasma metabolome of fattening Holstein steers fed a high-grain diet
Dhungana, A., Odunfa, O. A., Altman, A., Ogunade, I., McLeod, K. R., Yoon, I., Harmon, D. L. & Jiang, Y., 2025, In: Journal of Animal Science. 103, skaf307.Research output: Contribution to journal › Article › peer-review
Gastrointestinal development and tight junctions of preweaned calves supplemented with sodium butyrate in the starter mixture and whole milk
Zhang, Z., Wu, D., Li, X., Jia, Y., Qi, J., Harmon, D. L. & Xu, M., Jun 2025, In: Tropical Animal Health and Production. 57, 5, 234.Research output: Contribution to journal › Article › peer-review
In vitro antimicrobial efficacy of Cannabis sativa L. (hemp) crude oil extract on Fusobacterium necrophorum of bovine origin
Lakes, J. E., Altman, A. W., Berhow, M. A., Kagan, I. A., Nagaraja, T. G., Harmon, D. L. & Flythe, M. D., Jun 2025, In: Anaerobe. 93, 102960.Research output: Contribution to journal › Article › peer-review
Nutrient net absorption across the portal-drained viscera of forage-fed beef steers: Quantitative assessment and application to a nutritional prediction model
El-Kadi, S. W., McLeod, K. R., Elam, N. A., Kitts, S. E., Taylor, C. C., Harmon, D. L., Bequette, B. J. & Vanzant, E. S., Sep 2008, In: Journal of Animal Science. 86, 9, p. 2277-2287 11 p.Research output: Contribution to journal › Article › peer-review
Whole stillage inclusion level influences in vitro fiber digestibility and ruminal fermentation of tall fescue hay
Ajayi, H. J., Davis, B. E., Lehmkuhler, J. W., Harmon, D. L., Jiang, Y. & Trotta, R. J., 2025, In: Journal of Animal Science. 103, skaf006.Research output: Contribution to journal › Article › peer-review
Influence of flake density and starch retrogradation on in vitro gas production kinetics, digestibility, and ruminal fermentation characteristics of steam-flaked corn
Trotta, R. J., Kreikemeier, K. K. & Harmon, D. L., 2024, In: Journal of Animal Science. 102, skae308.Research output: Contribution to journal › Article › peer-review
Influence of postruminal casein infusion and exogenous glucagon-like peptide 2 administration on the jejunal mucosal transcriptome in cattle
Trotta, R. J., Swanson, K. C., Klotz, J. L. & Harmon, D. L., Aug 1 2024, In: PLoS ONE. 19, 8, e0308983.Research output: Contribution to journal › Article › peer-review
Influence of rearing method on the behaviour and welfare of Assaf lambs during the neonatal period
Simeonov, M., Harmon, D. L. & Stoycheva, I., Sep 2024, In: Indian Journal of Animal Sciences. 94, 9, p. 797-802 6 p.Research output: Contribution to journal › Article › peer-review
Effects of a Proprietary Kelp Blend Product on Enteric Methane Production and Tissue Residues in Cattle
Altman, A., Vanzant, E., Davis, S., Harmon, D. & McLeod, K. R., Aug 2024, In: Animals. 14, 16, 2411.Research output: Contribution to journal › Article › peer-review
Levodopa attenuates the feed intake reduction caused by ergot alkaloids in cattle
Valente, E. E. L., Klotz, J. L., Markmann, R. C., Trotta, R. J., Edwards, J. L., May, J. B. & Harmon, D. L., 2024, In: Journal of Animal Science. 102, skae078.Research output: Contribution to journal › Article › peer-review
Impact of endophyte-infected tall fescue seed consumption on endocrine changes associated with intake regulation and post-absorptive metabolism in growing steers
King, M. E., Herzing, H. M., McLeod, K. R., Klotz, J. L., Foote, A. P., Edwards, J. L. & Harmon, D. L., Oct 2024, In: Domestic Animal Endocrinology. 89, 106873.Research output: Contribution to journal › Article › peer-review
Serotonin receptor-mediated vasorelaxation occurs primarily through 5-HT4 activation in bovine lateral saphenous vein
Trotta, R. J., Harmon, D. L. & Klotz, J. L., Jul 2024, In: Physiological Reports. 12, 13, e16128.Research output: Contribution to journal › Article › peer-review
Nutritional and Physiological Constraints Contributing to Limitations in Small Intestinal Starch Digestion and Glucose Absorption in Ruminants
Trotta, R. J., Harmon, D. L., Matthews, J. C. & Swanson, K. C., Mar 2022, In: Ruminants. 2, 1, p. 1-26 26 p.Research output: Contribution to journal › Review article › peer-review
5-hydroxytryphophan mitigates ergot alkaloid-induced suppression of serotonin and feed intake in cattle
Valente, E. E. L., Klotz, J. L., Markmann, R. C., Edwards, J. L. & Harmon, D. L., 2024, In: Journal of Animal Science. 102, skae083.Research output: Contribution to journal › Article › peer-review
Influence of α-linked glucose on sodium-glucose cotransport activity along the small intestine in cattle
Bauer, M. L., Harmon, D. L., Bohnert, D. W., Branco, A. F. & Huntington, G. B., Jul 2001, In: Journal of Animal Science. 79, 7, p. 1917-1924 8 p.Research output: Contribution to journal › Article › peer-review
Review: Utilizing industrial hemp (Cannabis sativa L.) by-products in livestock rations
Altman, A. W., Kent-Dennis, C., Klotz, J. L., McLeod, K. R., Vanzant, E. S. & Harmon, D. L., Jan 2024, In: Animal Feed Science and Technology. 307, 115850.Research output: Contribution to journal › Review article › peer-review
Exploring the Role of G Protein Expression in Sodium Butyrate-Enhanced Pancreas Development of Dairy Calves: A Proteomic Perspective
Wu, D., Zhang, Z., Wang, X., Harmon, D. L., Jia, Y., Qi, J., Li, X., Jia, H. & Xu, M., Mar 20 2024, In: Journal of Agricultural and Food Chemistry. 72, 11, p. 5645-5658 14 p.Research output: Contribution to journal › Article › peer-review
Effects of exogenous phytase on small intestinal phosphorus disappearance in cattle
Trotta, R. (PI) & Harmon, D. (CoI)
9/1/25 → 3/31/27
Project: Research project
ARS/FAPRU: Improving Sustainability of Forage-Based Production Systems
Harmon, D. (PI)
8/28/25 → 8/27/26
Project: Research project
The Agronomy of Hemp and Its Uses in Forage-Animal Agriculture
Harmon, D. (PI)
8/1/25 → 7/31/26
Project: Research project
ARS/FAPRU: Improving Sustainability of Forage-Based Production Systems
Matthews, J. (PI), Bridges, P. (CoI), Ely, D. (CoI), Haan, N. (CoI), Harmon, D. (CoI), Hunt, A. (CoI), Ji, H. (CoI), McCulley, R. (CoI), McLeod, K. (CoI), Phillips, T. (CoI), Schardl, C. (CoI), Smith, S. (CoI), Suman, S. (CoI), Teutsch, C. (CoI), Trotta, R. (CoI), VanValin, K. (CoI) & Zhu, H. (CoI)
8/28/25 → 8/27/26
Project: Research project
The Agronomy of Hemp and Its Uses in Forage-Animal Agriculture
Matthews, J. (PI), Gauthier, N. (CoI), Harmon, D. (CoI), Hunt, A. (CoI), Ji, H. (CoI), Kawashima, T. (CoI), Lawrence, L. (CoI), Mark, T. (CoI), Moe, L. (CoI), Page, A. (CoI), Pearce, R. (CoI), Schendel, R. (CoI), Villanueva, R. (CoI) & Xiong, Y. (CoI)
8/1/25 → 7/31/26
Project: Research project
Rapid evaluation of exogenous glucoamylases on blood glucose appearance from starch digestion in cattle
Trotta, R. (PI) & Harmon, D. (CoI)
5/29/25 → 3/31/27
Project: Research project
ARS/FAPRU: Improving Sustainability of Forage-Based Production Systems
Matthews, J. (PI), Anderson, L. (CoI), Bridges, P. (CoI), Ely, D. (CoI), Haan, N. (CoI), Harmon, D. (CoI), Hunt, A. (CoI), Ji, H. (CoI), McCulley, R. (CoI), McLeod, K. (CoI), Phillips, T. (CoI), Rentfrow, G. (CoI), Schardl, C. (CoI), Smith, S. (CoI), Suman, S. (CoI), Teutsch, C. (CoI), VanValin, K. (CoI) & Zhu, H. (CoI)
5/1/23 → 4/30/26
Project: Research project
ARS/FAPRU: Harmon - Improving Sustainability of Forage-Based Production Systems
Harmon, D. (PI)
5/1/23 → 4/30/26
Project: Research project
Potential Amelioration Of Fescue Toxicosis By 5-Hydroxytryptophan In Beef Cattle
Trotta, R. (PI) & Harmon, D. (CoI)
National Institute of Food and Agriculture
8/1/24 → 7/31/26
Project: Research project
Mitigation of Sustained Ergot Alkaloid-Induced Vasoconstriction in Beef Cattle by Serotonin
Trotta, R. (PI) & Harmon, D. (CoI)
National Institute of Food and Agriculture
5/24/24 → 5/24/24
Project: Research project
The Agronomy of Hemp and Its Uses in Forage-Animal Agriculture
Harmon, D. (PI), Ely, D. (CoI), McLeod, K. (CoI) & Vanzant, E. (CoI)
8/1/21 → 7/31/26
Project: Research project
The Agronomy of Hemp and Its Uses in Forage-Animal Agriculture
Matthews, J. (PI), Adam, E. (CoI), Adams, A. (CoI), Ely, D. (CoI), Gauthier, N. (CoI), Harmon, D. (CoI), Hunt, A. (CoI), Ji, H. (CoI), Kawashima, T. (CoI), Lawrence, L. (CoI), Mark, T. (CoI), McCulley, R. (CoI), McLeod, K. (CoI), Moe, L. (CoI), Page, A. (CoI), Pearce, R. (CoI), Schendel, R. (CoI), Urschel, K. (CoI), Vanzant, E. (CoI), Villanueva, R. (CoI) & Xiong, Y. (CoI)
8/1/21 → 7/31/26
Project: Research project
Effects of Exogenous Glucoamylase on Small Intestine Starch Disappearance on Cattle
Harmon, D. (PI)
4/1/23 → 3/31/25
Project: Research project
Assessment of Available Literature and Gap Analysis on the Use of Industrial Hemp as a Livestock Feed
Harmon, D. (PI)
Foundation for Food and Agriculture Research
11/1/22 → 10/31/23
Project: Research project
Impact of Feeding Foods Containing Industrial Hemp on Health and Well-Being of Dogs
Harmon, D. (PI) & McLeod, K. (CoI)
2/27/18 → 8/26/21
Project: Research project
Services Agreement - Rumen Cannulation of Cows
Harmon, D. (PI)
4/5/18 → 12/31/18
Project: Research project
Student Support Agreement - Amanda Pesqueira
Harmon, D. (PI)
8/1/16 → 5/31/22
Project: Research project
Establishment of GLP quality systems for UK Animal Science
Harmon, D. (PI)
7/1/15 → 2/6/17
Project: Research project